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fills - you select a brush, you select trees, you draw on the map, and then you click "fill." The area you selected will be filled with trees of the type you chose in the menu.
regions - creating regions for scripts. Only for them.
masks - a block of the cell or cells that are highlighted by masks. It will not be possible to pass through it, even if it is completely open, etc.
All of this is in the manuals, which have already been translated into Russian.