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Nope, relative to the ground. Air cannot be taken as a reference frame. Imagine the wind is blowing. Relative to the airflow, the plane is moving, but not relative to the ground.
The plane accelerates due to the turbines, but it moves relative to the ground, and its speed increases, so its momentum also increases. But on the runway, the plane's speed will be 0, which means its momentum will also be zero.
And in general, your statement, "No matter how hard we try, their speeds (wheels and runway) will never be the same," contradicts the condition. Because we are told that the speeds are equal.